Yun, Goro and their robot Jet Jaguar clash with a hostile pterosaur-like kaiju, Rodan. Mei travels to Tokyo to meet with a world-renowned researcher.

A legion of Rodan swarm towards Nigashio. The Otaki Factory crew scramble to lure them away from the city, but soon find themselves surrounded.

Yun and Haberu deduce that Misakioku may be linked to the Rodan incursion. Mei attends Li's lecture in Dubai, where the professor reveals new material.

Salunga emerges from the depths of the Upala research station shrouded in red dust. Meanwhile, Yun observes Anguirus exhibiting an uncanny ability.

Mei discovers a mysterious code in Professor Ashihara's research. The Otaki crew confront Anguirus as part of the city's extermination efforts.

Following a group of Manda, a giant kaiju makes landfall in Tokyo. Red dust forces Mei and Professor Li's flight to London to divert to Milan.

Ground forces under the command of Matsubara respond to Godzilla's attack on Tokyo. Now in London, Mei attempts to decipher Ashihara's research.

Godzilla continues to advance on Tokyo, covering the city in red dust. The Otaki crew rushes to rescue a person ensnared by spider-like kaiju.

Back at Ashihara's mansion, Yun and Haberu attempt to decipher the professor's research using Jet Jaguar. In India, Lina takes Mei to meet with BB.

Before heading to the Super Dimension Calculator, BB offers Mei a view of the Singular Point. The Otaki crew leave for Tokyo to destroy Godzilla.

Mei learns of a large explosion triggered by Ashihara's calculator long ago. Kanoko gives Matsubara the Orthogonal Diagonalizer to use against Godzilla.

Mei, BB and Pelops II attempt to prevent the Catastrophe, but Salunga ambushes them. Meanwhile, the Otaki crew make their final push toward Godzilla.